Really interesting question...
@Chris Thomson
I remember there was a bit of underwater footage on the Barbel Days & Ways DVDs of a group of Barbel having a good grub about in a very silty marginal spot, with practically zero flow.
Generally speaking, it would be the last place you'd expect to find them, according to the 'textbooks', but there they were!
Just to caveat that, albeit with a single scenario, there's a quite well known holding spot on the Royalty that has filled with exactly the type of sandy substrate that you referenced. This occurred in last winter's floods. The depth, weed growth, etc hasn't drastically changed. It's just silty sand where there was once clean gravel. As far as I'm aware, it hasn't produced much more than a few juveniles this season. But I've caught in the next swim down and know of a few fish coming out in the next available swim upstream.
It'll be really interesting to see if this Winter's high water on the HAvon (assuming we get it for the same prolonged period, as is par for the course these days), clears the silty areas that are forming or exacerbates them... And if the swim in question does clear, whether it returns to its previous form!